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Comprehensive knowledge of the distribution of active hydrothermal vent fields along midocean ridges is essential to understanding global chemical and heat fluxes and endemic faunal distributions. However, current knowledge is biased by a historical preference for on-axis surveys. A scarcity of high-resolution bathymetric surveys in off-axis regions limits vent identification, which implies that the number of vents may be underestimated. Here, we present the discovery of an active, high-temperature, off-axis hydrothermal field on a fast-spreading ridge. The vent field is located 750 m east of the East Pacific Rise axis and ∼7 km north of on-axis vents at 9° 50′N, which are situated in a 50- to 100-m-wide trough. This site is currently the largest vent field known on the East Pacific Rise between 9 and 10° N. Its proximity to a normal fault suggests that hydrothermal fluid pathways are tectonically controlled. Geochemical evidence reveals deep fluid circulation to depths only 160 m above the axial magma lens. Relative to on-axis vents at 9° 50′N, these off-axis fluids attain higher temperatures and pressures. This tectonically controlled vent field may therefore exhibit greater stability in fluid composition, in contrast to more dynamic, dike-controlled, on-axis vents. The location of this site indicates that high-temperature convective circulation cells extend to greater distances off axis than previously realized. Thorough high-resolution mapping is necessary to understand the distribution, frequency, and physical controls on active off-axis vent fields so that their contribution to global heat and chemical fluxes and role in metacommunity dynamics can be determined. 

The Cook-Austral volcanic lineament extends from Macdonald Seamount (east) to Aitutaki Island (west) in the South Pacific Ocean and consists of hotspot-related volcanic islands, seamounts, and atolls. The Cook-Austral volcanic lineament has been characterized as multiple overlapping, age-progressive hotspot tracks generated by at least two mantle plumes, including the Arago and Macdonald plumes, which have fed volcano construction for ~20 m.y. The Arago and Macdonald hotspot tracks are argued to have been active for at least 70 m.y. and to extend northwest of the Cook-Austral volcanic lineament into the Cretaceous-aged Tuvalu-Gilbert and Tokelau Island chains, respectively. Large gaps in sampling exist along the predicted hotspot tracks, complicating efforts seeking to show that the Arago and Macdonald hotspots have been continuous, long-lived sources of hotspot volcanism back into the Cretaceous. We present new major- and trace-element concentrations and radiogenic isotopes for three seamounts (Moki, Malulu, Dino) and one atoll (Rose), and new clinopyroxene 40Ar/39Ar ages for Rose (24.81 ± 1.02 Ma) and Moki (44.53 ± 10.05 Ma). All volcanoes are located in the poorly sampled region between the younger Cook-Austral and the older, Cretaceous portions of the Arago and Macdonald hotspot tracks. Absolute plate motion modeling indicates that the Rose and Moki volcanoes lie on or near the reconstructed traces of the Arago and Macdonald hotspots, respectively, and the 40Ar/39Ar ages for Rose and Moki align with the predicted age progression for the Arago (Rose) and Macdonald (Moki) hotspots, thereby linking the younger Cook-Austral and older Cretaceous portions of the long-lived (>70 m.y.) Arago and Macdonald hotspot tracks.